Pinus hokkaidoensis

Name

Pinus hokkaidoensis

Rank

Species

Generic Name

[Genus] Pinus

Authors (Pub.)

Stockey R. A. Ueda Y.  

Publication

Permineralized pinaceous leaves from the Upper Cretaceous of Hokkaido [1986/8]

Journal

American Journal of Botany

Volume

73

Issue

8

Page number

1159

Year

1986

Fossil Status

foliage (needles)

Stratigraphy

Santonian

Strat. comment

Upper Yezo Group

Location

Minorizawa of the Sankebetsugawa, Haboro, Hokkaido, Japan

Paleoregion

Eurasia (Japanese Archipelago)

Data for Holotypus

Repository Number

832225

Num

Figs 3-7

Pub.

Permineralized pinaceous leaves from the Upper Cretaceous of Hokkaido [1986/8]

Diagnosis

Needles probably in fascicles of five, at least 0.5 cm long, 0.8-1.0 mm by 0.7-0.8 mm in tangential and radial widths respectively; fascicle sheath unknown. Vascular strand double, separated by a band of fibers one cell wide that expands on both adaxial and abaxial sides to four to five cells in radial thickness. Phloem not preserved; secondary xylem rows in each bundle 5-7 tracheids long. Transfusion tissue two to three cells thick; cell shapes irregular 35-63 ,m and 20-30 ,um in radial and tangential widths respectively. Endodermal cells large up to 65 ,m in diam with not appreciably thickened outer walls; no casparian strips visible. Resin canals two, external, located on adaxial sides of tangential corners. Mesophyll one cell thick, reaching two to three cells thick at corners; cells slightly plicate, 30-65 ,um in diam. Hypodermis uniform, one to four cells thick. Epidermal cells thick-walled circular to slightly elliptical in shape, identical to hypodermis. Stomata sunken; epidermal cells adjacent to stomatal opening beaked in transverse section; leaves amphistomatic.